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Brandt informs me that he has compared the hair from the face of a man thus characterised, aged thirty-five, with the lanugo of a foetus, and finds it quite similar in texture; therefore, as he remarks, the case may be attributed to an arrest of development in the hair, together with its continued growth. Brandt has recently sent me an additional case of a father and son, born in Russia, with these peculiarities.

I have received drawings of both from Paris. It appears as if the posterior molar or wisdom-teeth were tending to become rudimentary in the more civilised races of man. These teeth are rather smaller than the other molars, as is likewise the case with the corresponding teeth in the chimpanzee and orang; and they have only two separate fangs.

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They do not cut through the gums till about the seventeenth year, and I have been assured that they are much more liable to decay, and are earlier lost than the other teeth; but this is denied by some eminent dentists. They are also much more liable to vary, both in structure and in the period of their development, than the other teeth. Schaaffhausen accounts for this difference between the races by "the posterior dental portion of the jaw being always shortened" in those that are civilised,[44] and this shortening may, I presume, be attributed to civilised men habitually feeding on soft, cooked food, and thus using their jaws less.

I am informed by Mr. Brace that it is becoming quite a common practice in the United States to remove some of the molar teeth of children, as the jaw does not grow large enough for the perfect development of the normal number. The caecum is a branch or diverticulum of the intestine, ending in a cul-de-sac, and is extremely long in many of the lower vegetable-feeding mammals.

In the marsupial koala it is actually more than thrice as long as the whole body. Carter Blake in 'Anthropological Review,' July , p.

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Montegazza writes to me from Florence, that he has lately been studying the last molar teeth in the different races of man, and has come to the same conclusion as that given in my text, viz. That this appendage is a rudiment, we may infer from its small size, and from the evidence which Prof. Canestrini[47] has collected of its variability in man. It is occasionally quite absent, or again is largely developed. The passage is sometimes completely closed for half or two-thirds of its length, with theterminal part consisting of a flattened solid expansion.

In the orang this appendage is long and convoluted: in man it arises from the end of the short caecum, and is commonly from four to five inches in length, being only about the third of an inch in diameter. Not only is it useless, but it is sometimes the cause of death, of which fact I have lately heard two instances: this is due to small hard bodies, such as seeds, entering the passage, and causing inflammation. Now in the humerus of man, there is generally a trace of this passage, which is sometimes fairly well developed, being formed by a depending hook-like process of bone, completed by a band of ligament.

Struthers,[49] who has closely attended to the subject, has now shewn that this peculiarity is sometimes inherited, as it has occurred in a father, and in no less than four out of his seven children. When present, the great nerve invariably passes through it; and this clearly indicates that it is the homologue and rudiment of the supra-condyloid foramen of the lower animals.

Turner estimates, as he informs me, that it occurs in about one per cent. But if the occasional development of this structure in man is, as seems probable, due to reversion, it is a return to a very ancient state of things, because in the higher Quadrumana it is absent. Struthers in the 'Lancet,' Feb. Knox, as I am informed, was the first anatomist who drew attention to this peculiar structure in man; see his 'Great Artists and Anatomists,' p. See also an important memoir on this process by Dr. Gruber, in the 'Bulletin de l'Acad. This occurs, but not constantly, in various anthropoid and other apes,[50] and likewise in many of the lower animals.

It is remarkable that this perforation seems to have been present in man much more frequently during ancient times than recently. Busk[51] has collected the following evidence on this head: Prof. Broca "noticed the perforation in four and a half per cent.


Dupont found thirty per cent. Leguay, in a sort of dolmen at Argenteuil, observed twenty-five per cent. Pruner-Bey found twenty-six per cent. Nor should it be left unnoticed that M. Pruner-Bey states that this condition is common in Guanche skeletons. One chief cause seems to be that the ancient races stand somewhat nearer in the long line of descent to their remote animal-like progenitors. In man, the os coccyx, together with certain other vertebrae hereafter to be described, though functionless as a tail, plainly represent this part in other vertebrate animals.

At an early embryonic period it is free, and projects beyond the lower extremities; as may be seen in the drawing Fig. Even after birth it has been known, in certain rare and anomalous cases,[52] to form a small external rudiment of a tail. George Mivart, 'Transact. Congress of Prehist.

It frequently occurs in the negro. Turner, has been expressly described by Theile as a rudimentary repetition of the extensor of the tail, a muscle which is so largely developed in many mammals.

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The spinal cord in man extends only as far downwards as the last dorsal or first lumbar vertebra; but a thread-like structure the filum terminale runs down the axis of the sacral part of the spinal canal, and even along the back of the coccygeal bones. The upper part of this filament, as Prof. Turner informs me, is undoubtedly homologous with the spinalcord; but the lower part apparently consists merely of the pia mater, or vascular investing membrane.

Even in this case the os coccyx may be said to possess a vestige of so important a structure as the spinal cord, though no longer enclosed within a bony canal. The following fact, for which I am also indebted to Prof. Turner, shews how closely the os coccyx corresponds with the true tail in the lower animals: Luschka has recently discovered at the extremity of the coccygeal bones a very peculiar convoluted body, which is continuous with the middle sacral artery; and this discovery led Krause and Meyer to examine the tail of a monkey Macacus , and of a cat, in both of which they found a similarly convoluted body, though not at the extremity.

The reproductive system offers various rudimentary structures; but these differ in one important respect from the foregoing cases. Here we are not concerned with the vestige of a part which does not belong to the species in an efficient state, but with a part efficient in the one sex, and represented in the other by a mere rudiment. Nevertheless, the occurrence of such rudiments is as difficult to explain, on the belief of the separate creation of each species, as in the foregoing cases.

Hereafter I shall have to recur to these rudiments, and shall shew that their presence generally depends merely on inheritance, that is, on parts acquired by one sex having been partially transmitted to the other. I will in this place only give some instances of such rudiments. It is well known that in the males of all mammals, including man, rudimentary mammae exist. These in several instances have become well developed, and have yielded a copious supply of milk. The vesicula prostatica , which has been observed in many male mammals, is now universally acknowledged to be the homologue of the female uterus, together with the connected passage.

It is impossible to read Leuckart's able description of this organ, and his reasoning, without admitting the justness of his conclusion. This is especially clear in the case of those mammals in which the true female uterus bifurcates, for in the males of these the vesicula likewise bifurcates. But it would be superfluous fully to recapitulate the line of argument given in detail in my 'Origin of Species. In man this organ is only from three to six lines in length, but, like so many other rudimentary parts, it is variable in development as well as in other characters.

No one has shewn so well, how admirably such structures are adapted for their final purpose; and this adaptation can, as I believe, be explained through natural selection. In considering the wing of a bat, he brings forward p. It is unfortunate that he did not consider such cases as the minute teeth, which never cut through the jaw in the ox, or the mammae of male quadrupeds, or the wings of certain beetles, existing under the soldered wing-covers, or the vestiges of the pistil and stamens in various flowers, and many other such cases.

Although I greatly admire Prof. Bianconi's work, yet the belief now held by most naturalists seems to me left unshaken, that homological structures are inexplicable on the principle of mere adaptation. In order to understand the existence of rudimentary organs, we have only to suppose that a former progenitor possessed the parts in question in a perfect state, and that under changed habits of life they became greatly reduced, either from simple disuse, or through the natural selection of those individuals which were least encumbered with a superfluous part, aided by the other means previously indicated.

Thus we can understand how it has come to pass that man and all other vertebrate animals have been constructed on the same general model, why they pass through the same early stages of development, and why they retain certain rudiments in common.

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Consequently we ought frankly to admit their community of descent: to take any other view, is to admit that our own structure, and that of all the animals around us, is a mere snare laid to entrap our judgment. This conclusion is greatly strengthened, if we look to the members of the whole animal series, and consider the evidence derived from their affinities or classification, their geographical distribution and geological succession.

It is only our natural prejudice, and that arrogance which made our forefathers declare that they were descended from demi-gods, which leads us to demur to this conclusion. But the time will before long come, when it will be thought wonderful that naturalists, who were well acquainted with the comparative structure and development of man, and other mammals, should have believed that each was the work of a separate act of creation.

IT is manifest that man is now subject to much variability.

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No two individuals of the same race are quite alike. We may compare millions of faces, and each will be distinct. There is an equally great amount of diversity in the proportions and dimensions of the various parts of the body; the length of the legs being one of the most variable points. The chief arteries so frequently run in abnormal courses, that it has been found useful for surgical purposes to calculate from corpses how often each course prevails.

Turner[4] not to be strictly alike in any two out of fifty bodies; and in some the deviations were considerable. Statistics of American Soldiers,' by B. Gould, , p.

Aitken Meigs in 'Proc. On the Sandwich Islanders, Prof. Wyman, 'Observations on Crania,' Boston, , p. Preface, vol.